A
study conducted when Koko was 15 years old was designed
to see how accurately Koko understands spoken English,
specifically the smallest units of speech sound or phonemes.
Dr. Penny Patterson showed Koko sets of flash cards
with photos of various objects that differed by just
one phoneme, and then asked her for one of them by name.
For example, one set of cards required that Koko distinguish
between honey, money, bunny and funny. The results,
summarized below, were presented to the American Society
of Primatology in June, 2007.
As a subject in a longitudinal language acquisition
project, Koko communicates using sign language, which
she learned through the simultaneous communication of
spoken English and American Sign Language (ASL). To
test her understanding of spoken English she was presented
with pictures representing words taken from her vocabulary
(55 sets of 2, 3, or 4) that differed by one phoneme,
either vowel or consonant, in the initial, medial, or
final positions. Koko was able significantly to discriminate
phonemes in a linguistic context (78.2%). Koko's scores
were best when tested for final word sounds (100%) and
when offered 4 words per test item (90%). Her poorest
performance was with medial consonant word sounds when
offered only 2 words per item (55.6%).

These results suggest that motivational as well as perceptual
factors may have played a role in test performance.
Nevertheless this finding indicates phoneme discrimination
ability in a gorilla and supports the theory that speech
perception evolutionarily preceded speech production.
The presentation was during a session entitled Learning,
Cognition and Language, grouped with other talks related
to these headings. For example a talk in the same session
showed long-term memory capabilities of captive chimpanzees
who not only remembered the location of 20 different
hidden food rewards, but also prioritized the locations
by food quantity, type, and proximity. It was fascinating
to see what other researchers are doing to show the
amazing cognitive abilities of non-human primates. 
Our research focus at the next few conferences will
shift to promoting two-way communication with gorillas
and other non-human great apes as a form of enrichment
and a husbandry aid. We have been working on developing
technology to aid institutions who wish to expand on
the natural ability of apes to gesture by teaching them
to communicate with their keepers. Stay tuned for updates.
Dr. Christa Nunes
Associate Director of Research
(Note: The above presentation was given by Dr. Christa
Nunes, based on results collected by researchers: May
Goodreau, Christine Tam and Dr. Penny Patterson (Director
of Research).)
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